Monday, July 28, 2008

Amphibian reproduction

Now on reproduction of amphibians. For the purpose of reproduction most amphibians are bound to have fresh water. They need to be laid in damp places because of their jellylike characteristic and because there is no shell which means the eggs would quickly dry out and the young would die before they even had a chance to develop if not in water. When the eggs are laid, the male releases sperm over the female's eggs as they are laid. During this time, the male is in a posture called Amplexus which may be inguinal (male grasping the female in front of the her back legs) or axillary (male grasping the female in front of the her hands).
Several hundred frog species in adaptive radiations (e.g. Eleutherodactylus), however, do not need any water whatsoever. They reproduce via direct development, an ecological and evolutionary adaptation that has allowed them to be completely independent from free-standing water. Almost all of these frogs live in wet tropical rainforests and their eggs hatch directly into miniature versions of the adult, passing through the tadpole stage within the egg.
Several species have also adapted to arid and semi-arid environments, but most of them still need water to lay their eggs. Symbiosis with single celled algae that lives in the jelly-like layer of the eggs has evolved several times. The larvae (tadpoles or polliwogs) breathe with exterior gills.
Other kinds of fertilisation include those by salamanders whereby the male deposits a packet of sperm called a spermatophore onto the ground. The female then pulls it into her cloaca where fertilization occurs internally. Some caecilians and tailed frogs use internal fertilization where the male deposits sperm directly into the female's cloaca via an intromittent organ.
Amphibians laying their eggs in water normally leave them alone, but those laying eggs on land usually watch over them to protect them from predators. A few have developed strange ways to guard their eggs until they have hatched. The Surinam toad (Pipa pipa) of South America carries its eggs in pockets of skin on its back until the froglets hatch three months later. The gastric brooder (Rheobatrachus silus) of Australia no sooner lays its eggs than it swallows them! The eggs incubate in the frog's stomach where acid production ceases until the froglets hatch and crawl out through their mother's mouth.
Once hatched, the larva feed on yolk from their egg until they are able to find food for themselves. Then, they start to transform gradually into the adult's appearance. This process is called metamorphosis. Typically, the animals then leave the water and become terrestrial adults.
I thank http://www.eoearth.org/article/Amphibian_morphology_and_reproduction and wikipedia for the information.

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